In a thermostat, the R wire is responsible for powering the entire HVAC system (through a transformer). If you have both an Rh (Red-heat for powering the heating) and an Rc (Red-cool for powering the cooling), then the R wire can be connected to either Rh or Rc. If you only have a single R wire, then it will only have an R connector, which is the connection you’ll use.
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